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The Black Death probably originated in Central Asia or East Asia, from where it travelled along the Silk Road, reaching Crimea by 1347. From there, it was most likely carried by fleas living on the black rats that travelled on Genoese merchant ships, spreading throughout the Mediterranean Basin and reaching Africa, Western Asia, and the rest of Europe via Constantinople, Sicily, and the Italian Peninsula.
The Black Death is estimated to have killed 30% to 60% of Europe's population. In total, the plague may have reduced the world population from an estimated 475 million to 350–375 million in the 14t century. It took 200 years for Europe's population to recover to its previous level. Outbreaks of the plague recurred at various locations around the world until the early 20th century.
Recent research has suggested plague first infected humans in Europe and Asia in the Late Neolithic-Early Bronze Age.[32] Research in 2018 found evidence of Yersinia pestis in an ancient Swedish tomb, which may have been associated with the "Neolithic decline" around 3000 BC, in which European populations fell significantly. This Y. pestis may have been different to more modern types, with bubonic plague transmissible by fleas first known from Bronze Age remains near Samara.
The symptoms of bubonic plague are first attested in a fragment of Rufus of Ephesus preserved by Oribasius; these ancient medical authorities suggest bubonic plague had appeared in the Roman Empire before the reign of Trajan, six centuries before arriving at Pelusium in the reign of Justinian I. In 2013, researchers confirmed earlier speculation that the cause of the Plague of Justinian (541–542 AD, with recurrences until 750) was Yersinia pestis.This is known as the First Plague pandemic.
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